Who does Twilly directly share his feelings of anger and frustration with about ecological loss?

Twilly's direct sharing of his feelings of anger and frustration regarding ecological loss occurs with Nick and Martha. This interaction emphasizes the importance of personal relationships in dealing with complex emotions about environmental issues. By confiding in Nick and Martha, Twilly finds a space to articulate his concerns and frustrations, demonstrating the role that close friends or confidants play in navigating feelings related to significant societal problems. Their responses and support may lead to deeper discussions about ecological loss and the impact it has on the community, thereby reinforcing the themes of connection and activism within the narrative.